The Biota of the Gordon Natural Area - Bryophytes
Bryophytes (non-vascular plants which include the mosses, liverworts, and hornworts) have received relatively little investigation at the GNA. In 2007, WCU undergrad Spencer Ruston, under the supervision of Dr. Sharon Began, conducted a site-wide survey of bryophytes at the GNA, in which he noted 17 species. In 2009, undergrad Cindy Law, also under the supervision of Dr. Began, began another site-wide study of the GNA’s bryophytes. Although she didn’t finish this work, she did compile a photographic record of the eight species of bryophytes that she encountered. No additional work was done on bryophytes from that time until 2019, at which point GNA staff began photographing and collecting bryophytes with an aim towards create a more complete account of this biotic group at the GNA. The principal staff associated with this work are GNA Stewardship Manager Nur Ritter and GNA Student Intern Noah Long.
To date, 36 species of bryophytes, in 29 Genera, 23 Families, and 13 Orders have been noted at the GNA. This number is thought to significantly under-represent bryophyte richness at the GNA. Currently, we have numerous yet-to-be-identified species captured in photographs, and we hope to be able to use these to augment the species list. Because much of the bryophyte work at the GNA has employed photographs, this biotic group is extremely well-represented in our photo collection, with photos available for nearly nine-tenths (86.2%) of the known GNA species.
Note: the checklist is also available as a pdf
Aneura pinguis (Small Greasewort)
Group: Bryophytes
Type: Liverworts
Family: Aneuraceae
Notes
Common Name: Small Greasewort
Year Noted: 2020-2021
Identified By: Scott Schuette (iNaturalist
First Noted By: Nur Ritter
iNaturalist Observations:
66627530 (Observed: 12/15/2020)
52527487 (Observed: 7/9/2020)
89855001 (Observed: 8/3/2021)
References:
Lincoln, M. S. G. (2008). Liverworts of New England: A Guide for the Amateur Naturalist. Bronx, NY, New York Botanical Garden Press (p. 37*; 38).
Pope, R. Mosses, Liverworts, and Hornworts: A Field Guide to Common Bryophytes of the Northeast. Ithaca, NY, Cornell University Press (p. 299*).
Note: An asterisk after a page number signifies that: 1) the species is only mentioned on that page; or, 2) only the genus is discussed on that page.
Images
Frullania eboracensis (New York Scalewort)
Group: Bryophytes
Type: Liverworts
Family: Frullaniaceae
Notes
Common Names: New York Scalewort; Common Scalewort
Years Noted: 2017; 2019-2026
Identified and First Noted By: Nur Ritter
Also Noted By: Noah Long; Maeve Dowd
Comments:
Seemingly, the most common leafy liverwort at the Gordon. Observed growing on the bark of Northern Red Oak (Quercus rubra), White Ash (Fraxinus americana), Tulip Poplar (Liriodendron tulipifera), and Red Maple (Acer rubrum).
iNaturalist Observations:
326478707 (Observed: 11/13/2025)
99537627 (Observed: 10/26/2021)
37590246 (Observed: 1/13/2020)
50602427 (Observed: 6/22/2020)
References:
Conard, H. S. (1944). How to Know the Mosses. Mt. Pleasant, Iowa, H. E. Jacques (p. 131).
Klips, R. (2022). Common Mosses, Liverworts, and Lichens of Ohio. Columbus, OH, Ohio University Press (p. 212).
Davison, P. G. and M. J. Pistrang (2008). A Trailside Guide to Mosses and Liverworts of the Cherokee National Forest, Published by the authors using Blurb Inc. (p. 14).
Lincoln, M. S. G. (2008). Liverworts of New England: A Guide for the Amateur Naturalist. Bronx, NY, New York Botanical Garden Press (p. 120).
Pope, R. Mosses, Liverworts, and Hornworts: A Field Guide to Common Bryophytes of the Northeast. Ithaca, NY, Cornell University Press (p. 331).
Images
Lophocolea bidentata (Bifid Crestwort)
Group: Bryophytes
Type: Liverworts
Family: Geocalycaceae
Notes
Common Name: Bifid Crestwort
Year Noted: 2021
Identified and First Noted By: Nur Ritter
iNaturalist Observations:
87897234 (Observed: 7/20/2021)
References:
Conard, H. S. (1944). How to Know the Mosses. Mt. Pleasant, Iowa, H. E. Jacques (p. 122).
Lincoln, M. S. G. (2008). Liverworts of New England: A Guide for the Amateur Naturalist. Bronx, NY, New York Botanical Garden Press (p. 78).
Lophocolea heterophylla (Variable-leaved Crestwort)
Group: Bryophytes
Type: Liverworts
Family: Geocalycaceae
Notes
Common Name: Variable-leaved Crestwort
Years Noted: 2007; 2009; 2019-2023; 2025
Identified and First Noted By: Spencer Ruston
Also Noted By: Cindy Law; Nur Ritter; Paige Vermeulen; Kathryn Krueger
Comments:
Quite common at the GNA. This species and Frullania eboracensis are the two most commonly encountered Liverworts at the GNA.
iNaturalist Observations:
146893656 (Observed: 1/19/2023)
138613712 (Observed: 10/11/2022)
106746665 (Observed: 2/11/2022)
105617465 (Observed: 1/25/2022)
89831887 (Observed: 8/3/2021)
39765233 (Observed: 3/9/2020)
40103246 (Observed: 3/16/2020)
42226802 (Observed: 4/14/2020)
66390184 (Observed: 12/10/2020)
66627891 (Observed: 12/15/2020)
68614117 (Observed: 1/25/2021)
References:
Conard, H. S. (1944). How to Know the Mosses. Mt. Pleasant, Iowa, H. E. Jacques (p. 122).
Lincoln, M. S. G. (2008). Liverworts of New England: A Guide for the Amateur Naturalist. Bronx, NY, New York Botanical Garden Press (p. 77).
Munch, S. (2006). Outstanding Mosses and Liverworts of Pennsylvania and Nearby States. New Kingston, PA, Sunbury Press (p. 77).
Pope, R. Mosses, Liverworts, and Hornworts: A Field Guide to Common Bryophytes of the Northeast. Ithaca, NY, Cornell University Press (p. 336).
Images
Frullania inflata (Inflated Scalewort)
Group: Bryophytes
Type: Liverworts
Family: Jubulaceae
Notes
Common Name: Inflated Scalewort
Year Noted: 2026
Identified and First Noted By: Nur Ritter
iNaturalist Observations:
335743236 (Observed: 1/22/2026)
References:
Lincoln, M. S. G. (2008). Liverworts of New England: A Guide for the Amateur Naturalist. Bronx, NY, New York Botanical Garden Press (p. 125*).
Note: An asterisk after a page number signifies that: 1) the species is only mentioned on that page; or, 2) only the genus is discussed on that page.
Cololejeunea biddlecomiae (Biddlecomes Pouncewort)
Group: Bryophytes
Type: Liverworts
Family: Lejeuneaceae
Notes
Common Name: Biddlecome's Pouncewort
Year Noted: 2025-2026
Identified and First Noted By: Nur Ritter
iNaturalist Observations:
330374790 (Observed: 12/11/2025)
References:
Lincoln, M. S. G. (2008). Liverworts of New England: A Guide for the Amateur Naturalist. Bronx, NY, New York Botanical Garden Press (p. 115).
Pallavicinia lyellii (Veilwort)
Group: Bryophytes
Type: Liverworts
Family: Pallaviciniaceae
Notes
Common Names: Veilwort; Ribbonwort
Years Noted: 2021-2024; 2026
Identified and First Noted By: Nur Ritter
iNaturalist Observations:
334004790 (Observed: 1/8/2026)
92523355 (Observed: 8/25/2021)
144417214 (Observed: 12/14/2022)
106747241 (Observed: 2/11/2022)
92513649 (Observed: 8/25/2021)
References:
Davison, P. G. and M. J. Pistrang (2008). A Trailside Guide to Mosses and Liverworts of the Cherokee National Forest, Published by the authors using Blurb Inc. (p. 30*).
Lincoln, M. S. G. (2008). Liverworts of New England: A Guide for the Amateur Naturalist. Bronx, NY, New York Botanical Garden Press (p. 35).
McKnight, K. B., et al. (2013). Common Mosses of the Northeast and Appalachians (Princeton Field Guides Book 86). Princeton, NJ, Princeton University Press (p. 313).
Munch, S. (2006). Outstanding Mosses and Liverworts of Pennsylvania and Nearby States. New Kingston, PA, Sunbury Press (p. 73).
Note: An asterisk after a page number signifies that: 1) the species is only mentioned on that page; or, 2) only the genus is discussed on that page.
Images
Porella pinnata (Pinnate Scalewort)
Group: Bryophytes
Type: Liverworts
Family: Porellaceae
Notes
Common Name: Pinnate Scalewort
Year Noted: 2025-2026
Identified By: Ryan Sorrells (iNaturalist)
First Noted By: Nur Ritter
iNaturalist Observations:
330343498 (Observed: 12/10/2025)
References:
Conard, H. S. (1944). How to Know the Mosses. Mt. Pleasant, Iowa, H. E. Jacques (p. 129).
Lincoln, M. S. G. (2008). Liverworts of New England: A Guide for the Amateur Naturalist. Bronx, NY, New York Botanical Garden Press (p. 128).
Munch, S. (2006). Outstanding Mosses and Liverworts of Pennsylvania and Nearby States. New Kingston, PA, Sunbury Press (p. 81).
Radula cf. complanata (Flatleaf Scalewort)
Group: Bryophytes
Type: Liverworts
Family: Radulaceae
Notes
Common Name: Flatleaf Scalewort
Year Noted: 2020
Identified and First Noted By: Nur Ritter
iNaturalist Observations:
42130702 (Observed: 4/9/2020)
References:
Conard, H. S. (1944). How to Know the Mosses. Mt. Pleasant, Iowa, H. E. Jacques (p. 131).
Lincoln, M. S. G. (2008). Liverworts of New England: A Guide for the Amateur Naturalist. Bronx, NY, New York Botanical Garden Press (p. 118).
Pope, R. Mosses, Liverworts, and Hornworts: A Field Guide to Common Bryophytes of the Northeast. Ithaca, NY, Cornell University Press (p. 346).
Images
Amblystegiaceae Indet. #1 (No Common Name)
Hygroamblystegium varium (Tangled Thread Moss)
Group: Bryophytes
Type: Mosses
Family: Amblystegiaceae
Notes
Common Name: Tangled Thread Moss
Year Noted: 2025
Identified and First Noted By: Nur Ritter
iNaturalist Observations:
325884835 (Observed: 11/4/2025)
References:
McKnight, K. B., et al. (2013). Common Mosses of the Northeast and Appalachians (Princeton Field Guides Book 86). Princeton, NJ, Princeton University Press (p. 211).
Anomodon attenuatus (Poodle Moss)
Group: Bryophytes
Type: Mosses
Family: Anomodontaceae
Notes
Common Names: Poodle Moss; Tree-skirt Moss
Years Noted: 2019-2020; 2022
Identified and First Noted By: Nur Ritter
iNaturalist Observations:
41189361 (Observed: 4/23/2019)
41858638 (Observed: 4/10/2020)
References:
McKnight, K. B., et al. (2013). Common Mosses of the Northeast and Appalachians (Princeton Field Guides Book 86). Princeton, NJ, Princeton University Press (p. 239).
Munch, S. (2006). Outstanding Mosses and Liverworts of Pennsylvania and Nearby States. New Kingston, PA, Sunbury Press (p. 47).
Ohio Moss & Lichens Association website (https://ohiomosslichen.org/moss-anomodon-attenuatus)
Pope, R. Mosses, Liverworts, and Hornworts: A Field Guide to Common Bryophytes of the Northeast. Ithaca, NY, Cornell University Press (p. 233).
Images
Brachytheciaceae Indet. #1 (No Common Name)
Group: Bryophytes
Type: Mosses
Family: Brachytheciaceae
Notes
Common Name: No Common Name
Year Noted: 2019
Identified By: Carlos J. Pasiche Lisboa (iNaturalist)
First Noted By: Nur Ritter
iNaturalist Observations:
38852686 (Observed: 5/29/2019)
Images
Bryoandersonia illecebra (Spoon-leaved Moss)
Group: Bryophytes
Type: Mosses
Family: Brachytheciaceae
Notes
Common Names: Spoon-leaved Moss; Worm Moss
Years Noted: 2007; 2019-2020; 2022; 2025
Identified and First Noted By: Spencer Ruston
Also Noted By: Nur Ritter; kojo1019 (iNaturalist)
iNaturalist Observations:
328239285 (Observed: 11/25/2025)
39533318 (Observed: 2/25/2020)
141697674 (Observed: 11/11/2022)
References:
McKnight, K. B., et al. (2013). Common Mosses of the Northeast and Appalachians (Princeton Field Guides Book 86). Princeton, NJ, Princeton University Press (p. 255).
Munch, S. (2006). Outstanding Mosses and Liverworts of Pennsylvania and Nearby States. New Kingston, PA, Sunbury Press (p. 47).
Ohio Moss & Lichens Association website (https://ohiomosslichen.org/moss-bryoandersonia-illecebra)
Pope, R. Mosses, Liverworts, and Hornworts: A Field Guide to Common Bryophytes of the Northeast. Ithaca, NY, Cornell University Press (p. 242).
Images
Rhynchostegium serrulatum (Steerecleus Moss)
Group: Bryophytes
Type: Mosses
Family: Brachytheciaceae
Notes
Common Names: Steerecleus Moss; Dark Beaked Moss; Beaked Comb Moss
Year Noted: 2025
Identified and First Noted By: Nur Ritter
iNaturalist Observations:
328235140 (Observed: 11/25/2025)
References:
McKnight, K. B., et al. (2013). Common Mosses of the Northeast and Appalachians (Princeton Field Guides Book 86). Princeton, NJ, Princeton University Press (p. 243).
Ohio Moss & Lichens Association website (https://ohiomosslichen.org/moss-rhynchostegium-serrulatum)
Climacium dendroides (Northern Tree Moss)
Group: Bryophytes
Type: Mosses
Family: Climaciaceae
Notes
Common Names: Northern Tree Moss; Palm Tree Moss; Tree Moss
Year Noted: 2019-2021
Identified and First Noted By: Nur Ritter
Also Noted By: Ariana Rivellini; Noah Long
iNaturalist Observations:
37914840 (Observed: 10/18/2019)
References:
McKnight, K. B., et al. (2013). Common Mosses of the Northeast and Appalachians (Princeton Field Guides Book 86). Princeton, NJ, Princeton University Press (p. 203).
Munch, S. (2006). Outstanding Mosses and Liverworts of Pennsylvania and Nearby States. New Kingston, PA, Sunbury Press (p. 49).
Pope, R. Mosses, Liverworts, and Hornworts: A Field Guide to Common Bryophytes of the Northeast. Ithaca, NY, Cornell University Press (p. 245*).
Note: An asterisk after a page number signifies that: 1) the species is only mentioned on that page; or, 2) only the genus is discussed on that page.
Images
Dichodontium pellucidum (Translucent Fork Moss)
Group: Bryophytes
Type: Mosses
Family: Dicranaceae
Notes
Common Name: Translucent Fork Moss
Year Noted: 2025
Identified and First Noted By: Nur Ritter
iNaturalist Observations:
301013993 (Observed: 7/25/2025)
Dicranella heteromalla (Fine Hair Moss)
Group: Bryophytes
Type: Mosses
Family: Dicranaceae
Notes
Common Names: Fine Hair Moss; Silky Forklet Moss
Years Noted: 2007; 2020
Identified and First Noted By: Spencer Ruston
Also Noted By: Nur Ritter
References:
McKnight, K. B., et al. (2013). Common Mosses of the Northeast and Appalachians (Princeton Field Guides Book 86). Princeton, NJ, Princeton University Press (p. 37).
Ohio Moss & Lichens Association website (https://ohiomosslichen.org/moss-dicranella-heteromalla)
Pope, R. Mosses, Liverworts, and Hornworts: A Field Guide to Common Bryophytes of the Northeast. Ithaca, NY, Cornell University Press (p. 122).
Images
Dicranum scoparium (Broom Moss)
Group: Bryophytes
Type: Mosses
Family: Dicranaceae
Notes
Common Names: Broom Moss; Wind-swept Moss
Year Noted: 2021
Identified and First Noted By: Nur Ritter
iNaturalist Observations:
77514249 (Observed: 5/5/2021)
References:
McKnight, K. B., et al. (2013). Common Mosses of the Northeast and Appalachians (Princeton Field Guides Book 86). Princeton, NJ, Princeton University Press (p. 47).
Munch, S. (2006). Outstanding Mosses and Liverworts of Pennsylvania and Nearby States. New Kingston, PA, Sunbury Press (p. 33).
Ohio Moss & Lichens Association website (https://ohiomosslichen.org/moss-dicranum-scoparium)
Pope, R. Mosses, Liverworts, and Hornworts: A Field Guide to Common Bryophytes of the Northeast. Ithaca, NY, Cornell University Press (p. 128).
Images
Ceratodon purpureus (Redshank)
Group: Bryophytes
Type: Mosses
Family: Ditrichaceae
Notes
Common Names: Redshank; Purple Moss; Red Roof Moss
Year Noted: 2022
Identified and First Noted By: Nur Ritter
iNaturalist Observations:
139258140 (Observed: 10/17/2022)
References:
McKnight, K. B., et al. (2013). Common Mosses of the Northeast and Appalachians (Princeton Field Guides Book 86). Princeton, NJ, Princeton University Press (p. 87).
Ohio Moss & Lichens Association website (https://ohiomosslichen.org/moss-ceratodon-purpureus)
Pope, R. Mosses, Liverworts, and Hornworts: A Field Guide to Common Bryophytes of the Northeast. Ithaca, NY, Cornell University Press (p. 133).
Entodon seductrix (Seductive Entodon Moss)
Group: Bryophytes
Type: Mosses
Family: Entodontaceae
Notes
Common Names: Seductive Entodon Moss; Round-stem Entodon Moss; Cord Glaze Moss
Years Noted: 2020; 2025-2026
Identified and First Noted By: Nur Ritter
iNaturalist Observations:
330367060 (Observed: 12/11/2025)
40985840 (Observed: 3/29/2020)
References:
McKnight, K. B., et al. (2013). Common Mosses of the Northeast and Appalachians (Princeton Field Guides Book 86). Princeton, NJ, Princeton University Press (p. 291).
Ohio Moss & Lichens Association website (https://ohiomosslichen.org/moss-entodon-seductrix)
Pope, R. Mosses, Liverworts, and Hornworts: A Field Guide to Common Bryophytes of the Northeast. Ithaca, NY, Cornell University Press (p. 242*).
Note: An asterisk after a page number signifies that: 1) the species is only mentioned on that page; or, 2) only the genus is discussed on that page.
Images
Fissidens adianthoides (Maidenhair Pocket Moss)
Group: Bryophytes
Type: Mosses
Family: Fissidentaceae
Notes
Common Names: Maidenhair Pocket Moss; Maidenhair Split-toothed Moss
Years Noted: 2009; 2019-2020
Identified and First Noted By: Cindy Law
Also Noted By: Nur Ritter; Ariana Rivellini
Comments:
Very common in the GNA in a variety of habitats.
References:
Conard, H. S. (1944). How to Know the Mosses. Mt. Pleasant, Iowa, H. E. Jacques (p. 32).
McKnight, K. B., et al. (2013). Common Mosses of the Northeast and Appalachians (Princeton Field Guides Book 86). Princeton, NJ, Princeton University Press (p. 169).
Munch, S. (2006). Outstanding Mosses and Liverworts of Pennsylvania and Nearby States. New Kingston, PA, Sunbury Press (p. 37).
Ohio Moss & Lichens Association website (https://ohiomosslichen.org/moss-fissidens-adianthoides)
Pope, R. Mosses, Liverworts, and Hornworts: A Field Guide to Common Bryophytes of the Northeast. Ithaca, NY, Cornell University Press (p. 137).
Images
Fissidens osmundoides (Fern Pocket Moss)
Group: Bryophytes
Type: Mosses
Family: Fissidentaceae
Notes
Common Names: Fern Pocket Moss; Osmund Fissidens Moss
Year Noted: 2019-2020
Identified and First Noted By: Nur Ritter
iNaturalist Observations:
45205291 (Observed: 5/7/2020)
50587451 (Observed: 6/22/2020)
References:
McKnight, K. B., et al. (2013). Common Mosses of the Northeast and Appalachians (Princeton Field Guides Book 86). Princeton, NJ, Princeton University Press (p. 167).
Images
Funaria Indet. #1 (No Common Name)
Group: Bryophytes
Type: Mosses
Family: Funariaceae
Notes
Common Name: No Common Name
Year Noted: 2007
Identified and First Noted By: Spencer Ruston
Physcomitrium pyriforme (Goblet Moss)
Group: Bryophytes
Type: Mosses
Family: Funariaceae
Notes
Common Names: Goblet Moss; Common Bladder Moss; Pear-shaped Urn Moss
Year Noted: 2020-2022
Identified and First Noted By: Nur Ritter
iNaturalist Observations:
110836307 (Observed: 4/8/2022)
77512449 (Observed: 5/5/2021)
40982646 (Observed: 3/29/2020)
42996320 (Observed: 4/23/2020)
46764631 (Observed: 5/21/2020)
References:
McKnight, K. B., et al. (2013). Common Mosses of the Northeast and Appalachians (Princeton Field Guides Book 86). Princeton, NJ, Princeton University Press (p. 151).
Ohio Moss & Lichens Association website (https://ohiomosslichen.org/moss-physcomitrium-pyriforme)
Images
Grimmia Indet. #1 (No Common Name)
Group: Bryophytes
Type: Mosses
Family: Grimmiaceae
Notes
Common Name: No Common Name
Year Noted: 2007
Identified and First Noted By: Spencer Ruston
Schistidium apocarpum Group (Schistidium Moss)
Group: Bryophytes
Type: Mosses
Family: Grimmiaceae
Notes
Common Names: Schistidium Moss; Prickly Cannikin Moss
Year Noted: 2019-2020
Identified and First Noted By: Nur Ritter
Comments:
Noted growing on boulders in a few areas of the Gordon.
iNaturalist Observations:
141701789 (Observed: 3/18/2019)
References:
McKnight, K. B., et al. (2013). Common Mosses of the Northeast and Appalachians (Princeton Field Guides Book 86). Princeton, NJ, Princeton University Press (p. 111).
Munch, S. (2006). Outstanding Mosses and Liverworts of Pennsylvania and Nearby States. New Kingston, PA, Sunbury Press (p. 55*).
Ohio Moss & Lichens Association website (https://ohiomosslichen.org/moss-schistidium-apocarpum)
Pope, R. Mosses, Liverworts, and Hornworts: A Field Guide to Common Bryophytes of the Northeast. Ithaca, NY, Cornell University Press (p. 151).
Note: An asterisk after a page number signifies that: 1) the species is only mentioned on that page; or, 2) only the genus is discussed on that page.
Images
Hedwigia ciliata (White-tipped Moss)
Group: Bryophytes
Type: Mosses
Family: Hedwigiaceae
Notes
Common Names: White-tipped Moss; Medusa Moss; Ciliate Hedwigia Moss
Years Noted: 2007; 2020
Identified and First Noted By: Spencer Ruston
Also Noted By: Nur Ritter
iNaturalist Observations:
42997585 (Observed: 4/23/2020)
References:
Conard, H. S. (1944). How to Know the Mosses. Mt. Pleasant, Iowa, H. E. Jacques (p. 37).
McKnight, K. B., et al. (2013). Common Mosses of the Northeast and Appalachians (Princeton Field Guides Book 86). Princeton, NJ, Princeton University Press (p. 275).
Munch, S. (2006). Outstanding Mosses and Liverworts of Pennsylvania and Nearby States. New Kingston, PA, Sunbury Press (p. 55).
Pope, R. Mosses, Liverworts, and Hornworts: A Field Guide to Common Bryophytes of the Northeast. Ithaca, NY, Cornell University Press (p. 251; 213).
Images
Hypnum imponens (Brocade Moss)
Group: Bryophytes
Type: Mosses
Family: Hypnaceae
Notes
Common Names: Brocade Moss; Feather Moss; Pellucid Plait Moss
Years Noted: 2007; 2019-2022; 2025
Identified and First Noted By: Spencer Ruston
Also Noted By: Nur Ritter; Kathryn Krueger
iNaturalist Observations:
139863944 (Observed: 10/21/2022)
102732320 (Observed: 12/7/2021)
41190648 (Observed: 4/26/2019)
66430335 (Observed: 12/10/2020)
References:
McKnight, K. B., et al. (2013). Common Mosses of the Northeast and Appalachians (Princeton Field Guides Book 86). Princeton, NJ, Princeton University Press (p. 181).
Munch, S. (2006). Outstanding Mosses and Liverworts of Pennsylvania and Nearby States. New Kingston, PA, Sunbury Press (p. 59).
Ohio Moss & Lichens Association website (https://ohiomosslichen.org/moss-hypnum-imponens)
Pope, R. Mosses, Liverworts, and Hornworts: A Field Guide to Common Bryophytes of the Northeast. Ithaca, NY, Cornell University Press (p. 266).
Images
Platygyrium repens (Oil Spill Moss)
Group: Bryophytes
Type: Mosses
Family: Hypnaceae
Notes
Common Names: Oil Spill Moss; Flat Brocade Moss; Platygyrium Moss
Years Noted: 2007; 2009-2010; 2019
Identified and First Noted By: Spencer Ruston
Also Noted By: Cindy Law; Gerry Hertel; Nur Ritter
Comments:
A very common species at the GNA.
References:
Conard, H. S. (1944). How to Know the Mosses. Mt. Pleasant, Iowa, H. E. Jacques (p. 94).
McKnight, K. B., et al. (2013). Common Mosses of the Northeast and Appalachians (Princeton Field Guides Book 86). Princeton, NJ, Princeton University Press (p. 223).
Ohio Moss & Lichens Association website (https://ohiomosslichen.org/moss-platygyrium-repens)
Pope, R. Mosses, Liverworts, and Hornworts: A Field Guide to Common Bryophytes of the Northeast. Ithaca, NY, Cornell University Press (p. 269).
Images
Forsstroemia trichomitria (Fan Moss)
Group: Bryophytes
Type: Mosses
Family: Leptodontaceae
Notes
Common Name: Fan Moss
Year Noted: 2022
Identified By: mjpapay (iNaturalist)
First Noted By: Nur Ritter
iNaturalist Observations:
123263604 (Observed: 6/24/2022)
References:
Conard, H. S. (1944). How to Know the Mosses. Mt. Pleasant, Iowa, H. E. Jacques (p. 78).
Davison, P. G. and M. J. Pistrang (2008). A Trailside Guide to Mosses and Liverworts of the Cherokee National Forest, Published by the authors using Blurb Inc. (p. 68).
McKnight, K. B., et al. (2013). Common Mosses of the Northeast and Appalachians (Princeton Field Guides Book 86). Princeton, NJ, Princeton University Press (p. 235).
Pope, R. Mosses, Liverworts, and Hornworts: A Field Guide to Common Bryophytes of the Northeast. Ithaca, NY, Cornell University Press (p. 276).
Images
Leucobryum albidum (White Moss)
Group: Bryophytes
Type: Mosses
Family: Leucobryaceae
Notes
Common Name: White Moss
Year Noted: 2007
Identified and First Noted By: Spencer Ruston
References:
McKnight, K. B., et al. (2013). Common Mosses of the Northeast and Appalachians (Princeton Field Guides Book 86). Princeton, NJ, Princeton University Press (p. 123).
Pope, R. Mosses, Liverworts, and Hornworts: A Field Guide to Common Bryophytes of the Northeast. Ithaca, NY, Cornell University Press (p. 154).
Leucobryum glaucum (Pincushion Moss)
Group: Bryophytes
Type: Mosses
Family: Leucobryaceae
Notes
Common Name: Pincushion Moss
Year Noted: 2019
Identified and First Noted By: Nur Ritter
iNaturalist Observations:
37822549 (Observed: 8/27/2019)
69764003 (Observed: 8/27/2019)
References:
McKnight, K. B., et al. (2013). Common Mosses of the Northeast and Appalachians (Princeton Field Guides Book 86). Princeton, NJ, Princeton University Press (p. 123).
Ohio Moss & Lichens Association website (https://ohiomosslichen.org/moss-leucobryum-glaucum)
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Plagiomnium ciliare (Saber Tooth Moss)
Group: Bryophytes
Type: Mosses
Family: Mniaceae
Notes
Common Names: Saber Tooth Moss; Wavy-leaf Moss
Years Noted: 2007; 2009; 2019-2021; 2023; 2026
Identified and First Noted By: Spencer Ruston
Also Noted By: Cindy Law; Nur Ritter; Noah Long
iNaturalist Observations:
334005920 (Observed: 1/8/2026)
77668928 (Observed: 5/6/2021)
38845436 (Observed: 5/16/2019)
References:
McKnight, K. B., et al. (2013). Common Mosses of the Northeast and Appalachians (Princeton Field Guides Book 86). Princeton, NJ, Princeton University Press (p. 141).
Munch, S. (2006). Outstanding Mosses and Liverworts of Pennsylvania and Nearby States. New Kingston, PA, Sunbury Press (p. 41).
Ohio Moss & Lichens Association website (https://ohiomosslichen.org/moss-plagiomnium-ciliare)
Pope, R. Mosses, Liverworts, and Hornworts: A Field Guide to Common Bryophytes of the Northeast. Ithaca, NY, Cornell University Press (p. 159).
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Plagiomnium cuspidatum (Woodsy Thyme-Moss)
Group: Bryophytes
Type: Mosses
Family: Mniaceae
Notes
Common Names: Woodsy Thyme-Moss; Baby Tooth Moss; Woodsy Leafy Moss
Years Noted: 2007; 2019-2020; 2022; 2024-2025
Identified and First Noted By: Spencer Ruston
Also Noted By: Nur Ritter; Paige Vermeulen
iNaturalist Observations:
264118743 (Observed: 3/6/2025)
203094405 (Observed: 3/18/2024)
141704946 (Observed: 11/27/2019)
107430719 (Observed: 2/22/2022)
40992178 (Observed: 3/29/2020)
References:
Klips, R. (2022). Common Mosses, Liverworts, and Lichens of Ohio. Columbus, OH, Ohio University Press (p. 105).
McKnight, K. B., et al. (2013). Common Mosses of the Northeast and Appalachians (Princeton Field Guides Book 86). Princeton, NJ, Princeton University Press (p. 139).
Munch, S. (2006). Outstanding Mosses and Liverworts of Pennsylvania and Nearby States. New Kingston, PA, Sunbury Press (p. 41).
Ohio Moss & Lichens Association website (https://ohiomosslichen.org/moss-plagiomnium-cuspidatum)
Pope, R. Mosses, Liverworts, and Hornworts: A Field Guide to Common Bryophytes of the Northeast. Ithaca, NY, Cornell University Press (p. 159).
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Rhizomnium Indet. #1 (Leafy Mosses)
Rhizomnium punctatum (Dotted Thyme Moss)
Group: Bryophytes
Type: Mosses
Family: Mniaceae
Notes
Common Names: Dotted Thyme Moss; Red Penny Moss; Dotted Leafy Moss
Year Noted: 2019-2020
Identified and First Noted By: Nur Ritter
iNaturalist Observations:
42998262 (Observed: 4/23/2020)
References:
Conard, H. S. (1944). How to Know the Mosses. Mt. Pleasant, Iowa, H. E. Jacques (p. 62).
McKnight, K. B., et al. (2013). Common Mosses of the Northeast and Appalachians (Princeton Field Guides Book 86). Princeton, NJ, Princeton University Press (p. 143).
Munch, S. (2006). Outstanding Mosses and Liverworts of Pennsylvania and Nearby States. New Kingston, PA, Sunbury Press (p. 43).
Ohio Moss & Lichens Association website (https://ohiomosslichen.org/moss-rhizomnium-punctatum)
Pope, R. Mosses, Liverworts, and Hornworts: A Field Guide to Common Bryophytes of the Northeast. Ithaca, NY, Cornell University Press (p. 163).
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Orthotrichum anomalum (Anomalous Bristle-moss)
Group: Bryophytes
Type: Mosses
Family: Orthotrichaceae
Notes
Common Name: Anomalous Bristle-moss
Years Noted: 2020-2022; 2025
Identified and First Noted By: Nur Ritter
iNaturalist Observations:
260913764 (Observed: 2/7/2025)
104416073 (Observed: 1/4/2022)
References:
McKnight, K. B., et al. (2013). Common Mosses of the Northeast and Appalachians (Princeton Field Guides Book 86). Princeton, NJ, Princeton University Press (p. 107).
Munch, S. (2006). Outstanding Mosses and Liverworts of Pennsylvania and Nearby States. New Kingston, PA, Sunbury Press (p. 55).
Pope, R. Mosses, Liverworts, and Hornworts: A Field Guide to Common Bryophytes of the Northeast. Ithaca, NY, Cornell University Press (p. 167).
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Orthotrichum cf. stellatum (Stellate Orthotrichum Moss)
Group: Bryophytes
Type: Mosses
Family: Orthotrichaceae
Notes
Common Name: Stellate Orthotrichum Moss
Years Noted: 2019; 2021
Identified and First Noted By: Nur Ritter
iNaturalist Observations:
70342090 (Observed: 2/28/2021)
References:
McKnight, K. B., et al. (2013). Common Mosses of the Northeast and Appalachians (Princeton Field Guides Book 86). Princeton, NJ, Princeton University Press (p. 103).
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Orthotrichum strangulatum (Drummond Moss)
Group: Bryophytes
Type: Mosses
Family: Orthotrichaceae
Notes
Common Names: Drummond Moss; Strangulate Orthotrichum Moss; Shy Bristle Moss
Year Noted: 2019-2020
Identified and First Noted By: Nur Ritter
Comments:
Growing on boulders in the Gordon. Noted occasionally in the forest south of Stadium Road.
References:
McKnight, K. B., et al. (2013). Common Mosses of the Northeast and Appalachians (Princeton Field Guides Book 86). Princeton, NJ, Princeton University Press (p. 109).
Munch, S. (2006). Outstanding Mosses and Liverworts of Pennsylvania and Nearby States. New Kingston, PA, Sunbury Press (p. 55*).
Ohio Moss & Lichens Association website (https://ohiomosslichen.org/moss-orthotrichum-strangulatum)
Note: An asterisk after a page number signifies that: 1) the species is only mentioned on that page; or, 2) only the genus is discussed on that page.
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Atrichum altecristatum (Wavy Starburst Moss)
Group: Bryophytes
Type: Mosses
Family: Polytrichaceae
Notes
Common Name: Wavy Starburst Moss
Year Noted: 2019
Identified and First Noted By: Nur Ritter
Comments:
Seemingly, fairly common in the GNA. In shaded areas, often near streams.
References:
McKnight, K. B., et al. (2013). Common Mosses of the Northeast and Appalachians (Princeton Field Guides Book 86). Princeton, NJ, Princeton University Press (p. 69).
Ohio Moss & Lichens Association website (https://ohiomosslichen.org/moss-atrichum-altecristatum)
Images
Atrichum angustatum (Lesser Smoothcap Moss)
Group: Bryophytes
Type: Mosses
Family: Polytrichaceae
Notes
Common Names: Lesser Smoothcap Moss; Slender Starburst Moss
Years Noted: 2007; 2009; 2019; 2021
Identified and First Noted By: Spencer Ruston
Also Noted By: Cindy Law; Nur Ritter
iNaturalist Observations:
141699914 (Observed: 4/27/2021)
References:
McKnight, K. B., et al. (2013). Common Mosses of the Northeast and Appalachians (Princeton Field Guides Book 86). Princeton, NJ, Princeton University Press (p. 67).
Munch, S. (2006). Outstanding Mosses and Liverworts of Pennsylvania and Nearby States. New Kingston, PA, Sunbury Press (p. 21).
Ohio Moss & Lichens Association website (https://ohiomosslichen.org/moss-atrichum-angustatum)
Pope, R. Mosses, Liverworts, and Hornworts: A Field Guide to Common Bryophytes of the Northeast. Ithaca, NY, Cornell University Press (p. 173).
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Atrichum undulatum (Wavy Catharinea)
Group: Bryophytes
Type: Mosses
Family: Polytrichaceae
Notes
Common Name: Wavy Catharinea
Years Noted: 2007; 2019
Identified and First Noted By: Spencer Ruston
Also Noted By: Nur Ritter
References:
Conard, H. S. (1944). How to Know the Mosses. Mt. Pleasant, Iowa, H. E. Jacques (p. 27).
Munch, S. (2006). Outstanding Mosses and Liverworts of Pennsylvania and Nearby States. New Kingston, PA, Sunbury Press (p. 21).
Pope, R. Mosses, Liverworts, and Hornworts: A Field Guide to Common Bryophytes of the Northeast. Ithaca, NY, Cornell University Press (p. 175).
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Pogonatum pensilvanicum (Algal Haircap Moss)
Group: Bryophytes
Type: Mosses
Family: Polytrichaceae
Notes
Common Name: Algal Haircap Moss
Year Noted: 2025
Identified and First Noted By: Nur Ritter
iNaturalist Observations:
330365408 (Observed: 12/11/2025)
References:
McKnight, K. B., et al. (2013). Common Mosses of the Northeast and Appalachians (Princeton Field Guides Book 86). Princeton, NJ, Princeton University Press (p. 319).
Ohio Moss & Lichens Association website (https://ohiomosslichen.org/moss-pogonatum-pensylvanicum)
Polytrichum ohioense (Ohio Haircap Moss)
Group: Bryophytes
Type: Mosses
Family: Polytrichaceae
Notes
Common Names: Ohio Haircap Moss; Oak Haircap Moss
Years Noted: 2007; 2020
Identified and First Noted By: Spencer Ruston
Also Noted By: Nur Ritter
iNaturalist Observations:
38831947 (Observed: 2/18/2020)
References:
McKnight, K. B., et al. (2013). Common Mosses of the Northeast and Appalachians (Princeton Field Guides Book 86). Princeton, NJ, Princeton University Press (p. 59).
Ohio Moss & Lichens Association website (https://ohiomosslichen.org/moss-polytrichastrum-ohioense)
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Pottiaceae Indet. #1 (No Common Name)
Ptychomitrium incurvum (Incurved Grizzled Moss)
Group: Bryophytes
Type: Mosses
Family: Ptychomitriaceae
Notes
Common Names: Incurved Grizzled Moss; Gnome's Hood Moss
Years Noted: 2019; 2025-2026
Identified By: Carlos J. Pasiche Lisboa (iNaturalist)
First Noted By: Nur Ritter
iNaturalist Observations:
330253426 (Observed: 12/10/2025)
40749522 (Observed: 3/28/2019)
References:
McKnight, K. B., et al. (2013). Common Mosses of the Northeast and Appalachians (Princeton Field Guides Book 86). Princeton, NJ, Princeton University Press (p. 93).
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Sphagnum cf. fimbriatum (Fringed Bogmoss)
Group: Bryophytes
Type: Mosses
Family: Sphagnaceae
Notes
Common Names: Fringed Bogmoss; Fringed Peat Moss
Years Noted: 2020; 2025
Identified and First Noted By: Nur Ritter
iNaturalist Observations:
58979391 (Observed: 9/8/2020)
References:
McKnight, K. B., et al. (2013). Common Mosses of the Northeast and Appalachians (Princeton Field Guides Book 86). Princeton, NJ, Princeton University Press (p. 339).
Ohio Moss & Lichens Association website (https://ohiomosslichen.org/moss-sphagnum-fimbriatum)
Pope, R. Mosses, Liverworts, and Hornworts: A Field Guide to Common Bryophytes of the Northeast. Ithaca, NY, Cornell University Press (p. 40).
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Sphagnum cf. girgensohnii (Girgensohns Sphagnum)
Group: Bryophytes
Type: Mosses
Family: Sphagnaceae
Notes
Common Names: Girgensohn's Sphagnum; Graceful Peat Moss
Year Noted: 2025
Identified and First Noted By: Nur Ritter
iNaturalist Observations:
279711733 (Observed: 5/2/2025)
References:
McKnight, K. B., et al. (2013). Common Mosses of the Northeast and Appalachians (Princeton Field Guides Book 86). Princeton, NJ, Princeton University Press (p. 337).
Pope, R. Mosses, Liverworts, and Hornworts: A Field Guide to Common Bryophytes of the Northeast. Ithaca, NY, Cornell University Press (p. 43).
Thuidium allenii (Allens Thuidium Moss)
Group: Bryophytes
Type: Mosses
Family: Thuidiaceae
Notes
Common Names: Allen's Thuidium Moss; Fernmoss
Year Noted: 2007
Identified and First Noted By: Spencer Ruston
Thuidium delicatulum (Delicate Fern Moss)
Group: Bryophytes
Type: Mosses
Family: Thuidiaceae
Notes
Common Names: Delicate Fern Moss; Common Fern Moss; Delicate Thuidium Moss
Years Noted: 2009; 2019; 2021-2022
Identified and First Noted By: Cindy Law
Also Noted By: Nur Ritter; Kathryn Krueger
Comments:
Very common throughout the forested areas of the Gordon.
iNaturalist Observations:
139865642 (Observed: 10/21/2022)
100600165 (Observed: 11/3/2021)
37824588 (Observed: 12/31/2019)
References:
McKnight, K. B., et al. (2013). Common Mosses of the Northeast and Appalachians (Princeton Field Guides Book 86). Princeton, NJ, Princeton University Press (p. 271).
Munch, S. (2006). Outstanding Mosses and Liverworts of Pennsylvania and Nearby States. New Kingston, PA, Sunbury Press (p. 65).
Ohio Moss & Lichens Association website (https://ohiomosslichen.org/moss-thuidium-delicatulum)
Pope, R. Mosses, Liverworts, and Hornworts: A Field Guide to Common Bryophytes of the Northeast. Ithaca, NY, Cornell University Press (p. 290).








































































































































































